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What BIOS version do you have? If you haven’t upgrade BIOS is probably best to do so.
In order to upgrade BIOS is probably best to put the E4500 then once upgraded try the E7500
Find below a link with the latest BIOS update.
I downgraded BIOS from the latest version to the minimum required version 0484 (using the old CPU) and now everything works as intended.
I have no idea what the reason is but i don't realy care ... it's working and I'm a happy-happy panda!
I can only guess that for old motherboards the latest version of BIOS is not always the best.