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Your going to have to ask Lenovo, but I suspect the BIOS does not know how to configure this processor.
The hardware may or may not be able to support the processor in data paths and voltages since they are both 775, You would have to go look up the 2 processors specifications and check the Land Listing and Signal Descriptions section to see if all 775 pins are the same. Just because the socket is the same size, does guarantee that the processors will be compatible.
Their hw manual http://download.lenovo.com/ibmdl/pub/pc/pccbbs/thinkcentre_pdf/42y4652.pdf pg 129 list their supported processors, however they have not updated the manual since -- Fourth Edition (September 2007) andhe most current BIOS for the unit came out
Version: 2OKT48A Release Date: 2008/12/05
But the processor did not release until Q3 2010 it is highly unlikely they have support code for the newer processor in this old of a BIOS.
Product Name Intel® Pentium® Processor E5700 (2M Cache, 3.00 GHz, 800 MHz FSB) Intel® Core™2 Duo Processor E4300 (2M Cache, 1.80 GHz, 800 MHz FSB) Code Name Wolfdale Conroe Status Launched Launched Launch Date Q3'10 Q3'06 Processor Number E5700 E4300